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africana Agulhas Bank analysis andersoni appear Bairdoppilata Bate Bay BH-9 borehole Boomplaas Bythocypris Campanian Campanian-Maastrichtian cent climate compared correlation Cretaceous cytheracean Cytherella dense depth detailed Dingle distribution dominant dorsal Dutoitella element environmental environments evidence fact Factor fauna Figure Formation genera genus glacial grass Holotype Igoda important increase indicate individual internal interpretation late lateral length levels locality lower Maastrichtian material mean Mfolozi River moderate Needs Camp Nibela nibelaensis occurs ostracod assemblages outcrops palaeoecology pattern percentage period Pleistocene population possible preference present probably proportions range recorded relatively Remarks represented restricted Richards Bay BH-9 ridge rounded samples Santonian sequence shelf shows similar slope South south-east Africa species specimens stage strata suggested surface Table Total number types unit UNIV upper variation various vegetation Zululand