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abbreviated absence African Museum sample appear approach Beaufort West District bio-zone Kitching 1977 bone boss broad Broili & Schröder Broom canine caniniform process Cape Province Cistecephalus bio-zone Cluver & Hotton collection condition considered cover deep delicate dentary described developed Dicynodon Diictodon galeops distinctive distinguishing Dunedin Quaggafontein 82 elevated feature figures forms front genus given grossarthi holotype illustrations included indicated interorbital intertemporal region King known larger lateral Leeu Kloof 43 locality lower jaw margin maxilla maxillary rim median mentioned narrow nostrils noted orbital palate palatine parietals pattern pineal foramen pineal opening possible postorbitals premaxilla preparietal present preservation reduction references relative ring separate septomaxilla shape side Skull and lower skull length skull roof Smith snout South African Museum species specimens surrounded sutural Tapinocephalus bio-zone Kitching Toerien Tropidostoma-Endothiodon assemblage zone tuskless tusks variability variation Width