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abbreviated according articulate brachiopods association author's name authors bedding benthic assemblages bicuspidata bivalves Blinkberg Bokkeveld Group Boucot brachiopods capital collected communities consists containing cross-bedded cycles delta deposited described Devonian Devonian rocks differences DISTRIBUTARY diversity dominated Eastern environment environmental fauna figures gastropod homalonotid Howison's Poort illustrations inarticulate indicate infaunal interpreted intertidal Islands italics known letters Lingula localities lower major marine marine invertebrates mark Member Middle mould nearshore Note occur Orbiculoidea original placed plant fossils Plectonotus possible quartz arenites range recorded reduction references represent reworking salinity sandstones scientific sediment SEDIMENTARY FACIES separates sequence shales shallow siltstone similar Smith species specimens Stratigraphic subtidal suggest Tables Tankard thick thin Touws River Town trace fossils trilobite Tropidoleptus unit upper valve Wagen Drift Formation Weltevrede Formation Western Cape Witpoort Formation Witteberg Group yielded Zoophycos