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abbreviated according articulate brachiopods association author's name authors basal bedding benthic assemblages bivalves Blinkberg Bokkeveld Group Boucot capital collected communities consists containing cycles delta deposited described Devonian Devonian rocks differences displaying DISTRIBUTARY diversity dominated Eastern environment environmental fauna figures gastropod homalonotid Howison's Poort illustrations inarticulate indicate infaunal interpreted intertidal italics known letters Lingula localities lower major marine marine invertebrates mark Member Middle mould Museum Natural History nearshore Note occur original periods placed plant fossils Plectonotus possible quartz arenites range recognized recorded references represent reworking salinity sandstones scientific sediment SEDIMENTARY FACIES separates sequence shales shallow siltstone similar Smith South African species specimens Stratigraphic subtidal suggest Tables Tankard thick thin Touws River trace fossils trilobite unit upper upward-fining valve Wagen Drift Formation Weltevrede Formation Western Cape Witpoort Formation Witteberg Group yielded Zoophycos