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abbreviated articulate brachiopods Australospirifer barrier islands benthic assemblages BENTHIC COMMUNITIES Blinkberg Formation Bokkeveld Group Boucot British Museum Natural bryozoa Chonetes COMMUNITIES AND SEDIMENTARY cross-bedded deeper-water subtidal conditions delta deposited Devonian rocks environmental External mould facies analysis Falkland Islands fauna Formation near Touws Frasnian gastropod genus Givetian Grahamstown high-diversity holotype homalonotid trilobite horizontal lamination Howison's Poort inarticulate infaunal interpreted Leda Lingula lepta LOWER WITTEBERG GROUP Museum Natural History orbiculoid and linguloid Orbiculoidea overlain paratypes parentheses pedicle valve Plectonotus sp possible bryozoa punctuation salinity sandstones scientific name sediment SEDIMENTARY FACIES shoreline shouldered italics siltstone and shale Silurian and Devonian South African Museum specimens Stratigraphic STRATIGRAPHY suspension feeders SWARTRUGGENS FORMATION Tankard & Barwis taxa Theron thin tidal tidal-flat Touws River trace fossils Trigonoglossa Tropidoleptus sp Upper Devonian valve of Lingula valve of Tropidoleptus Wagen Drift Formation wash-over Weltevrede Formation Western Cape Witpoort Formation Witteberg Group represent yielded marine invertebrate