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abdominal adult Ammonites antennae apex apical appear Barnard base bearing beetles body body chamber brown brownish Cape coast Collected Collignon colour common Coniacian covered dark decaying December described developed DIEGO differs dorsal view dravidicum eggs fairly Family females figures four Gauthiericeras genus Grossouvre hatched Head Hoepen Holotype inner keel larvae lateral legs length lobe Locality mandible margin Matsumoto measure middle Museum narrow Natal nymphs observed occur ornament pairs pale Paratype Peroniceras Peroniceras Peroniceras Zuluiceras posterior present Pupa R.S. Pieter Faure referred ribs River rounded segment separated setae Shell shiny short showing side similar single slightly South African Sowerby species specimens spines stage suture Syntypes third tridorsatum tubercles umbilical UNIVERSITY usually valves van Hoepen ventral whorls wide Zone Zululand